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The Formation of Vegetable Mould through the Action of Worms

The Formation of Vegetable Mould through the Action of Worms

The Formation of Vegetable Mould Through the Action of Worms, with Observations on their Habits is an 1881 book by Charles Darwin on earthworms. It was his last scientific book, and was published shortly before his death . Exploring earthworm behaviour and ecology, it continued the theme common throughout his work that gradual changes over long periods of time can lead to large and sometimes surprising consequences. It was the first significant work on soil bioturbation, although that term was not used by Darwin .


Charles Darwin photo

“But some degree of intelligence appears, as we shall see in the next chapter, to be exhibited in this work,—a result which has surprised me more than anything else in regard to worms.”

Charles Darwin book The Formation of Vegetable Mould through the Action of Worms

Source: The Formation of Vegetable Mould through the Action of Worms (1881), Chapter 1: Habits of Worms, p. 35. http://darwin-online.org.uk/content/frameset?pageseq=50&itemID=F1357&viewtype=image

Charles Darwin photo

“When the pots containing two worms which had remained quite indifferent to the sound of the piano, were placed on this instrument, and the note C in the bass clef was struck, both instantly retreated into their burrows.”

Charles Darwin book The Formation of Vegetable Mould through the Action of Worms

Source: The Formation of Vegetable Mould through the Action of Worms (1881), Chapter 1: Habits of Worms, p. 28. http://darwin-online.org.uk/content/frameset?pageseq=43&itemID=F1357&viewtype=image

Charles Darwin photo

“In this case, therefore, the worms judged with a considerable degree of correctness how best to draw the withered leaves of this foreign plant into their burrows; notwithstanding that they had to depart from their usual habit of avoiding the foot-stalk.”

Charles Darwin book The Formation of Vegetable Mould through the Action of Worms

Source: The Formation of Vegetable Mould through the Action of Worms (1881), Chapter 2: Habits of Worms, p. 70. http://darwin-online.org.uk/content/frameset?pageseq=85&itemID=F1357&viewtype=image

Charles Darwin photo
Charles Darwin photo

“Worms seize leaves and other objects, not only to serve as food, but for plugging up the mouths of their burrows; and this is one of their strongest instincts.”

Charles Darwin book The Formation of Vegetable Mould through the Action of Worms

Source: The Formation of Vegetable Mould through the Action of Worms (1881), Chapter 2: Habits of Worms, p. 58. http://darwin-online.org.uk/content/frameset?pageseq=73&itemID=F1357&viewtype=image

Charles Darwin photo

“The subject may appear an insignificant one, but we shall see that it possesses some interest; and the maxim "de minimis lex non curat," does not apply to science.”

Charles Darwin book The Formation of Vegetable Mould through the Action of Worms

Introduction, p. 2. http://darwin-online.org.uk/content/frameset?pageseq=17&amp;itemID=F1357&amp;viewtype=image <br class="br">de minimis non curat lex - The law does not concern itself with trifles. <br class="br">The Formation of Vegetable Mould through the Action of Worms (1881)

Charles Darwin photo

“With respect to the function of the calciferous glands, it is probable that they primarily serve as organs of excretion, and secondarily as an aid to digestion.”

Charles Darwin book The Formation of Vegetable Mould through the Action of Worms

Source: The Formation of Vegetable Mould through the Action of Worms (1881), Chapter 1: Habits of Worms, p. 49. http://darwin-online.org.uk/content/frameset?pageseq=64&amp;itemID=F1357&amp;viewtype=image

Charles Darwin photo

“The comparison here implied between the actions of one of the higher animals and of one so low in the scale as an earth-worm, may appear far-fetched; for we thus attribute to the worm attention and some mental power, nevertheless I can see no reason to doubt the justice of the comparison.”

Charles Darwin book The Formation of Vegetable Mould through the Action of Worms

Source: The Formation of Vegetable Mould through the Action of Worms (1881), Chapter 1: Habits of Worms, pp. 24-25. http://darwin-online.org.uk/content/frameset?pageseq=39&amp;itemID=F1357&amp;viewtype=image

Charles Darwin photo

“Earth-worms abound in England in many different stations. Their castings may be seen in extraordinary numbers on commons and chalk-downs, so as almost to cover the whole surface, where the soil is poor and the grass short and thin.”

Charles Darwin book The Formation of Vegetable Mould through the Action of Worms

Source: The Formation of Vegetable Mould through the Action of Worms (1881), Chapter 1: Habits of Worms, p. 9. http://darwin-online.org.uk/content/frameset?pageseq=24&amp;itemID=F1357&amp;viewtype=image

Charles Darwin photo

“When a worm is suddenly illuminated and dashes like a rabbit into its burrow—to use the expression employed by a friend—we are at first led to look at the action as a reflex one.”

Charles Darwin book The Formation of Vegetable Mould through the Action of Worms

Source: The Formation of Vegetable Mould through the Action of Worms (1881), Chapter 1: Habits of Worms, p. 23. http://darwin-online.org.uk/content/frameset?pageseq=38&amp;itemID=F1357&amp;viewtype=image

Charles Darwin photo

“M. Perrier found that their exposure to the dry air of a room for only a single night was fatal to them. On the other hand he kept several large worms alive for nearly four months, completely submerged in water.”

Charles Darwin book The Formation of Vegetable Mould through the Action of Worms

Source: The Formation of Vegetable Mould through the Action of Worms (1881), Chapter 1: Habits of Worms, pp. 12-13 http://darwin-online.org.uk/content/frameset?pageseq=27&amp;itemID=F1357&amp;viewtype=image.

Charles Darwin photo

“Every morning during certain seasons of the year, the thrushes and blackbirds on all the lawns throughout the country draw out of their holes an astonishing number of worms; and this they could not do, unless they lay close to the surface.”

Charles Darwin book The Formation of Vegetable Mould through the Action of Worms

Source: The Formation of Vegetable Mould through the Action of Worms (1881), Chapter 1: Habits of Worms, p. 16. http://darwin-online.org.uk/content/frameset?pageseq=31&amp;itemID=F1357&amp;viewtype=image

Charles Darwin photo

“As it is certain that worms swallow many little stones, independently of those swallowed while excavating their burrows, it is probable that they serve, like mill-stones, to triturate their food.”

Charles Darwin book The Formation of Vegetable Mould through the Action of Worms

Source: The Formation of Vegetable Mould through the Action of Worms (1881), Chapter 1: Habits of Worms, p. 18. http://darwin-online.org.uk/content/frameset?pageseq=33&amp;itemID=F1357&amp;viewtype=image

Charles Darwin photo

“In the year 1837, a short paper was read by me before the Geological Society of London, "On the Formation of Mould," in which it was shown that small fragments of burnt marl, cinders, &c., which had been thickly strewed over the surface of several meadows, were found after a few years lying at the depth of some inches beneath the turf, but still forming a layer.”

Charles Darwin book The Formation of Vegetable Mould through the Action of Worms

Introduction, p. 3. http://darwin-online.org.uk/content/frameset?pageseq=18&amp;itemID=F1357&amp;viewtype=image <br class="br">The Formation of Vegetable Mould through the Action of Worms (1881) <br class="br">Source: &#x27;Transactions Geolog. Soc.&#x27; vol. v. p. 505. Read November 1, 1837.

Charles Darwin photo

“As I was led to keep in my study during many months worms in pots filled with earth, I became interested in them, and wished to learn how far they acted consciously, and how much mental power they displayed.”

Charles Darwin book The Formation of Vegetable Mould through the Action of Worms

Introduction, p. 2-3. http://darwin-online.org.uk/content/frameset?pageseq=17&amp;itemID=F1357&amp;viewtype=image <br class="br">The Formation of Vegetable Mould through the Action of Worms (1881)

Charles Darwin photo

“It may be presumed that all animals which feed on various substances possess the sense of taste, and this is certainly the case with worms.”

Charles Darwin book The Formation of Vegetable Mould through the Action of Worms

Source: The Formation of Vegetable Mould through the Action of Worms (1881), Chapter 1: Habits of Worms, p. 32. http://darwin-online.org.uk/content/frameset?pageseq=47&amp;itemID=F1357&amp;viewtype=image

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