“Later evolutionary theorists of linear progress had to advance the overtly physical and historical claim that an ancestral lineage of arthropods actually turned over to become the first vertebrates (for the classical statement of the inversion theory, see William Patten, The Grand Strategy of Evolution, 1920).”

—  Stephen Jay Gould, book Leonardo's Mountain of Clams and the Diet of Worms

Source: Leonardo's Mountain of Clams and the Diet of Worms (1998) "Brotherhood by Inversion", p. 327

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Stephen Jay Gould274
American evolutionary biologist 1941–2002

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“Gaskell… had to keep his stately soldiers upright and uniformly oriented… by crafting the brain and spinal cord from an arthropod digestive tube, while forming a completely new gut below. …Gaskell thought that his move would rescue the theory of linear progress, with its necessary transition of arthropod into vertebrate, from the absurdities of the old inversion theory.”

Stephen Jay Gould book Leonardo's Mountain of Clams and the Diet of Worms

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“Geoffrey also recognized that the opposite orientations of gut and nervous system posed a problem for his claim that insects and vertebrates represent different versions of the same archetypal animal - and he proposed the first account of the inversion theory to resolve this threat to unification. …Geoffroy pursued the… aim of establishing a "unity of type" that could generate both arthropods and vertebrates from the same basic blueprint. …The single grand design includes a gut in the middle and the main nerve cords somewhere on the periphery.”

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“How ironic. In order to avoid the "nutty" theory of inversion, Gaskell invented the even odder notion of stomachs turning into brains with new guts forming below. No wonder then, that later biologists cast a plague on both speculative houses and opted instead for the obvious alternative: arthropods and vertebrates do not share the same anatomical plan at all, but rather represent two separate evolutionary developments of similar complexity from a much simpler common ancestor that grew neither a discrete gut nor a central nerve cord.”

Stephen Jay Gould book Leonardo's Mountain of Clams and the Diet of Worms

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“Gaskell could not abide this indecorous version of his beloved linear progress theory. He could not bear to imagine that the grand procession from jellyfish to man, orchestrated by an ever-increasing mass of nervous tissue, once paused in its stately and orderly march toward human consciousness in order to execute a fancy little flip, a clever jig of inversion, just at the sublime and definitive moment of entrance into the vertebral home stretch.”

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