“Heredity. Given any organ in a parent and the same or any other organ in its offspring, the mathematical measure of heredity is the correlation of these organs for pairs of parent and offspring... The word organ here must be taken to include any characteristic which can be quantitatively measured.”

—  Karl Pearson

"Mathematical Contributions to the Theory of Evolution III: Regression, Heredity and Panmixia", Philosophical Transactions of the Royal Society, Series A, Vol. 187 (1896) p. 259.

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Karl Pearson 65
English mathematician and biometrician 1857–1936

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