“We may consequently state the fundamental theorem of Natural Selection in the form : The rate of increase in fitness of any organism at any time is equal to its genetic variance in fitness at that time.”

—  Ronald Fisher , book The Genetical Theory of Natural Selection

Defining the fundamental theorem of natural selection, Ch. 2, p. 35.
The Genetical Theory of Natural Selection (1930)

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Ronald Fisher 28
English statistician, evolutionary biologist, geneticist, a… 1890–1962

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